Andrea (Anne Hathaway) is an aspiring journalist recent out of Northwestern College. Regardless of her ridicule for the self-love of the style trade, she lands a job “one million women would kill for,” junior private assistant to Miranda Priestly (Meryl Streep), the icy editor-in-chief of Runway trend journal. Andy plans to place up with Miranda’s weird and humiliating remedy for one yr in hopes of getting a job as a reporter or author some place else.
At first, Andy fumbles together with her job and matches in poorly along with her gossipy, fashion-conscious co-workers, particularly Miranda’s senior assistant Emily Charlton (Emily Blunt). Nevertheless, with the assistance of artwork director Nigel (Stanley Tucci), who lends her designer garments, she steadily learns her obligations and begins to decorate extra stylishly to point out her effort and dedication to the place. She additionally meets enticing younger author Christian Thompson (Simon Baker), who presents to assist her along with her profession. As she spends growing quantities of time at Miranda’s beck and name, issues come up in her relationships along with her school associates and her live-in boyfriend Nate (Adrian Grenier), a chef working his manner up the profession ladder.
Miranda is impressed by Andy and permits her to be the one to carry the treasured “Guide,” a mock-up of the upcoming version, to her house, alongside together with her dry cleansing. She is given directions by Emily about the place to go away the gadgets and is advised to not communicate with anybody within the dwelling. Andy arrives at Miranda’s residence solely to find that the directions she obtained are imprecise. As she tries to determine what to do, Andy begins to panic. Miranda’s twins (Caroline and Cassidy, performed by Colleen and Suzanne Dengel, respectively) falsely inform her she will go away the guide on the high of the steps simply as Emily has achieved on many events. On the prime of the steps, Andy interrupts Miranda and her husband having an argument. Mortified, Andy leaves the ebook and runs out of the house.
The following day, Miranda tells her that she desires the brand new unpublished Harry Potter guide for her daughters and, if Andy can not discover a copy, she can be fired. Andy desperately makes an attempt to search out the e-book, almost offers up, however in the end obtains it by means of Christian’s contacts. She surprises Miranda by not solely discovering the e book however having copies despatched to the women on the prepare station, leaving little question that she completed Miranda’s “inconceivable” process, thus saving her job.
Someday, Andy saves Miranda from being embarrassed at a charity profit, and Miranda rewards her by providing to take her to the autumn style exhibits in Paris as an alternative of Emily. Andy hesitates to take this privilege away from Emily however is pressured to simply accept the supply after being informed by Miranda that she is going to lose her job if she declines. Andy tries to inform Emily on her option to work. Nonetheless, Emily is hit by a automobile and Andy has to interrupt the unhealthy information whereas visiting her within the hospital.
When Andy tells Nate she goes to Paris, he’s angered by her refusal to confess that she’s grow to be what she as soon as ridiculed, and so they break up. As soon as there, Miranda, with out make-up, opens as much as Andy in regards to the impact Miranda’s impending divorce may have on her daughters. Later that night time, Nigel tells Andy that he has accepted a job as Artistic Director with rising trend star James Holt (Daniel Sunjata) at Miranda’s advice. Andy lastly succumbs to Christian’s charms and, after spending the night time with him, learns from him a couple of plan to exchange Miranda with Jacqueline Follet as editor of Runway. Regardless of the struggling she has endured at her boss’s behest, she makes an attempt to warn Miranda.
At a luncheon later that day, nonetheless, Miranda pronounces that it’s Jacqueline as an alternative of Nigel who will go away Runway for Holt. Nigel remarks to a shocked Andy that, although upset, he has to imagine that his loyalty to Miranda will in the future repay. Later, when Miranda and Andy are being pushed to a present, she explains to a still-stunned Andy that she was grateful for the warning however already knew of the plot to exchange her and sacrificed Nigel to maintain her personal job. Happy by this show of loyalty, she tells Andy that she sees an excessive amount of herself in her. Andy, repulsed, says she might by no means try this to anybody. Miranda replies that she already did, stepping over Emily when she agreed to go to Paris. After they cease, Andy will get out and throws her mobile phone into the fountain of the Place de la Concorde, leaving Miranda, Runway, and style behind.
A while later, Andy is interviewed and is accepted to work at a significant New York publication firm. It was Miranda who insisted that they rent Andy, regardless of that she did not work for her a full 12 months, however the firm could be “the most important idiots” if they do not. After the interview, Andy meets up with Nate, who’s transferring to Boston as a result of he acquired a brand new job because the sous chef of a restaurant. They comply with begin relationship once more and see what the longer term holds. Andy calls Emily and affords her her Paris wardrobe and the 2 go away on good phrases. Andy passes the “Runway” workplace constructing and sees Miranda get right into a automotive. Andy provides a wave, however Miranda doesn’t acknowledge her. Andy is used to this and as a substitute walks additional into the group. As soon as contained in the automotive, nonetheless, Miranda smiles after which orders her chauffeur to drive.